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This companion article to interdigital perineural fibroma (a.k.a. Morton's neuroma [MN]) discusses the anatomy and physiology of the nerves in the forefoot, clinical presentation, diagnosis, and treatment of this painful fore foot condition. Although no empirical causal data exists to associate MN with Morton's syndrome or Morton's toe, a condition has been described in the literature whereby particular foot architecture has been linked to anomalies in the first and second metatarsals. This alteration in forefoot structure creates changes in foot biomechanics, precipitates foot pain, and alters foot functioning. To juxtapose this article with the possibility of Morton's syndrome, a family vignette referred to as "Jane's Genus" will be provided as an example of the prevalence of Morton's toe in one family.