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Abstract
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1 Japan Fisheries Research and Education Agency, Seikai National Fisheries Research Institute, Nagasaki, Japan
2 Japan Fisheries Research and Education Agency, Kamiura Station, National Research Institute of Aquaculture, Saiki, Japan
3 Japan Fisheries Research and Education Agency, National Research Institute of Aquaculture, Watarai, Japan (GRID:grid.410851.9) (ISNI:0000 0004 1764 1824)
4 Japan Fisheries Research and Education Agency, Kamiura Station, National Research Institute of Aquaculture, Saiki, Japan (GRID:grid.410851.9)
5 Japan Fisheries Research and Education Agency, Seikai National Fisheries Research Institute, Nagasaki, Japan (GRID:grid.410851.9)
6 Nagasaki University, Organization for Marine Science and Technology, Graduate School of Fisheries and Environmental Sciences, Nagasaki, Japan (GRID:grid.174567.6) (ISNI:0000 0000 8902 2273)
7 Ehime University, Nishiura Station, South Ehime Fisheries Research Center, Minamiuwa, Japan (GRID:grid.255464.4) (ISNI:0000 0001 1011 3808)
8 Japan Fisheries Research and Education Agency, National Research Institute of Fisheries Science, Yokohama, Japan (GRID:grid.410851.9) (ISNI:0000 0004 1764 1824)
9 Kindai University, Aquaculture Research Institute, Nishimuro, Japan (GRID:grid.258622.9) (ISNI:0000 0004 1936 9967)
10 Hiroshima University, Department of Mathematical and Life Sciences, Graduate School of Science, Higashi-Hiroshima, Japan (GRID:grid.257022.0) (ISNI:0000 0000 8711 3200)
11 Headquarters, Japan Fisheries Research and Education Agency, Yokohama, Japan (GRID:grid.410851.9) (ISNI:0000 0004 1764 1824)