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Abstract
The earliest stone tool types, sharp flakes knapped from stone cores, are assumed to have played a crucial role in human cognitive evolution. Flaked stone tools have been observed to be accidentally produced when wild monkeys use handheld stones as tools. Holding a stone core in hand and hitting it with another in the absence of flaking, free hand hitting, has been considered a requirement for producing sharp stone flakes by hitting stone on stone, free hand percussion. We report on five observations of free hand hitting behavior in two wild western gorillas, using stone-like objects (pieces of termite mound). Gorillas are therefore the second non-human lineage primate showing free-hand hitting behavior in the wild, and ours is the first report for free hand hitting behavior in wild apes. This study helps to shed light on the morphofunctional and cognitive requirements for the emergence of stone tool production as it shows that a prerequisite for free hand percussion (namely, free hand hitting) is part of the spontaneous behavioral repertoire of one of humans’ closest relatives (gorillas). However, the ability to combine free hand hitting with the force, precision, and accuracy needed to facilitate conchoidal fracture in free hand percussion may still have been a critical watershed for hominin evolution.
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1 Université Paris Cité, Eco-Anthropologie (EA), Muséum National d’Histoire Naturelle, CNRS, Paris, France (GRID:grid.508487.6) (ISNI:0000 0004 7885 7602)
2 UMR7179 MECADEV CNRS, Muséum National d’Histoire Naturelle, Department Adaptations du Vivant, Paris, France (GRID:grid.410350.3) (ISNI:0000 0001 2174 9334)
3 Wildlife Conservation Society, Bronx, USA (GRID:grid.269823.4) (ISNI:0000 0001 2164 6888); World Wide Fund for Nature – Germany, Berlin, Germany (GRID:grid.506609.c) (ISNI:0000 0001 1089 5299)
4 University of Tübingen, Department for Early Prehistory and Quaternary Ecology, Tübingen, Germany (GRID:grid.10392.39) (ISNI:0000 0001 2190 1447)